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How have we ruled out that successful mutations in the wild aren’t influenced by the environment?

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Sorry to repost this – just that I didn’t get an answer to my final question in this thread. Mostly, I think, because the question changed and because of my imprecise (or, at best, non technical) use of the world epigenetic. I now understand how we might be able to rule out successful adaptations in the lab being caused by survival-selected random mutations, but how have we ruled this out in the wild?

 

What evidence is there that successful adaptations in the wild are due to random mutations and not through some mechanism (genetic or otherwise) that responds to the environment?

 

Many thanks for your time.

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