I'm seeing all kinds of leptons, quarks, organisms, primates, experts, etc. on this site....I guess I'm wondering; whats the full hierarchical scale, and how does one advance?

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It's based on number of posts; you move up as you get more posts. (Posts in General Discussion and a few other non-sciency places don't count.)

 

Resident Experts were deemed to be science experts by the moderator and administrator staff because they exhibited expert knowledge of their subject on the forums. One of the perks of being a mod or admin is being able to choose a completely custom title, as you can see...